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Article

Pakasuchus kapilimai: Tanzania's Extinct Warm-Blooded Cat Crocodile

Tanzania’s fossils of extinct warm-blooded cat crocodile Pakasuchus kapilima discovered by Dr Patrick O’Connor head twenty-first century scientific finds.

Cat crocodiles do not bolt their food whole Among the scariest events in the 2008-released action drama horror film Black Water are the seconds during which a saltwater crocodile (Crocodylus porosus) achieves a vertical thrust of the upper body toward two Northern Australian swamp-touring sisters, Grace (Diana Glenn) and Lee (Maeve Dermody), whose feet dangle from low-lying mangrove branches. The terrifying upward movement brings to mind one of the unexpected constants in the life cycle and natural history of one of the saltwater crocodile’s distant ancestors, Pakasuchus kapilimai. Patrick O’Connor, one of the extinct antecedent’s discoverers and paleontologist of Ohio University at Athens, calls the specimens from East Africa’s Rukwa Rift Basin in southwestern Tanzania cat crocodiles, the literal meaning of Kiswahili paka and Greek suchus. ***** Website: http://www.oucom.ohiou.edu/dbms-oconnor/ *****

Ancient crocodile Pakasuchus kapilimai of Tanzania's Rukwa Rift Basin: graphic is comparing the size of the ancient crocodile Pakasuchus kapilimai to a housecat.

Pakasuchus was discovered by international researchers led by Patrick O'Connor of Ohio University and partly funded by NSF.
Pakasuchus was discovered by international researchers led by Patrick O'Connor of Ohio University and partly funded by NSF. — Public Domain, via Wikimedia Commons

Cat crocodiles exhibit mammal-like mouthparts for piecemeal chews

 

The description feline defers to such cat crocodile hallmarks as:

  • agile backbones;
  • housecat-sized 5- to 10-pound (2.27- to 4.54-kilogram) frames;
  • housecat-sized 2-foot (60.96-centimeter) head-to-tail lengths; and
  • small, 3.5-inch (8.89-centimeter) skulls.

The designation canine emphasizes a peculiarity of cat crocodile snouts whereby nasal openings are dog-like to the front for land-breathing as opposed to atop the head for water-breathing. The term mammal finds similar applicability regarding:

  • food chains and predator/prey webs;
  • look and use of jaws and teeth; and
  • niches within southern hemisphere habitats 65,000,000 to 105,000,000 years ago.

Paleontologists indeed give to cat crocodiles:

  • canine-, molar-, premolar-filled, mobile-jawed mouthparts;
  • diets of insects, primitive mammals, small lizards; and
  • roles of mammals with the break-up of the supercontinent Pangaea. 

 

Diagram of dentition of Early Cretaceous notosuchian Pakasuchus kapilimai. Gray area reconstructs missing parts of the holotype skull.

Pakasuchus kapilimai flourished as mammal-like Crocodyliforme in Cretaceous (Latin: creta, "chalk") geologic period.
Pakasuchus kapilimai flourished as mammal-like Crocodyliforme in Cretaceous (Latin: creta, "chalk") geologic period. — Smokeybjb, CC BY SA 3.0 (https://creativecommons.org/licenses/by-sa/3.0/deed.en), via Wikimedia Commons

Cat crocodiles favor insects, lizards, and primitive mammals

 

Southeast Africa’s floodplains hold for modern experts such as Professors Saidi Kapilima -- whom the cat crocodile species kapilimai honors -- of Tanzania’s University of Dar-es-Salaam and Patrick O’Connor of Ohio University at Athens red sandstone-preserved fossils from the southern supercontinent Gondwana. Medical scanning one such specimen’s tight-closed jaws indeed identifies the cat crocodile’s crocodilian skullbone arrangements, feline skull, and mammalian canines for biting, jaws and premolars for sliding, and molars for shearing before swallowing. Paleontologists thereby judge cat crocodile corporeal, dental, and mandibular forms and functions as consistent with the evolutionary precept of creatures pursuing similar lifestyles evolving similar features.

Fauna-loving amateurs, newbies, and specialists likewise know of the above-mentioned discoveries through Dr O’Connor’s taxonomy-related article of 2010. 

 

Pakasuchus kapilimai: an extinct, warm-blooded ancestor of saltwater crocodiles (Crocodilus porosus)

life restoration of Pakasuchus kapilimai
life restoration of Pakasuchus kapilimai — Smokeybjb, CC BY SA 3.0 (https://creativecommons.org/licenses/by-sa/3.0/deed.en), via Wikimedia Commons

Cat crocodiles get armor-free bodies with armor-full tails

 

Fossilized cat crocodiles let twenty-first century Earthlings in on ancient population controlling-species in mammal-sparse Gondwana to the south of animal-rich Laurasia. They make easier reconstructions of life cycles and natural histories regarding the collateral and direct ancestors of the twenty-first century’s fearsome alligators, caimans, crocodiles, and gavials. Their discovery indeed nudges the incredible and the unknown into less mysterious, more credible realms.

Cat crocodile fossils particularly offer insights into animal survival and faunal sustainability since the related crocodile and dinosaur descendants of archosaurs respectively did and did not survive ancient Earth’s mass extinctions. They preserve evidence of crocodilian experimentation with cat-like, dragonfly-chasing, front-snouted, land-lubbing, osteoderm-free, warm-blooded physiques hindered only by inexplicably weighted-down tails of bony, crocodile-like armor. 

 

"Pakasuchus" (1:10)

Uploaded August 8, 2010, by tyrantslayer24 to YouTube ~ URL: https://www.youtube.com/watch?v=DzJ3lxtJKFs

Pakasuchus

Acknowledgment

 

My special thanks to talented artists and photographers/concerned organizations who make their fine images available on the internet.

 

Image Credits

 

Ancient crocodile Pakasuchus kapilimai of Tanzania's Rukwa Rift Basin: graphic is comparing the size of the ancient crocodile Pakasuchus kapilimai to a housecat.
Roaming Tanzania in the Middle Cretaceous period, P. kapilimai had a number of features unusual for crocodylians, such as mammal-like teeth and flexible spine.The previously unknown "croc" was unearthed from 105 million-year-old, mid-Cretaceous rocks.
Pakasuchus was discovered by international researchers led by Patrick O'Connor of Ohio University and partly funded by NSF.: Public Domain, via Wikimedia Commons @ https://commons.wikimedia.org/wiki/File:Ancient_Crocodile_Pakasuchus_kapilimai_(15104697297).jpg; Zina Deretsky, National Science Foundation, Usage restrictions None, via EurekAlert! @ https://www.eurekalert.org/news-releases/733263 (specific image URL @  https://www.eurekalert.org/multimedia/894850)

Diagram of dentition of Early Cretaceous notosuchian Pakasuchus kapilimai. Gray area reconstructs missing parts of the holotype skull.
Pakasuchus kapilimai flourished as mammal-like Crocodyliforme in Cretaceous (Latin: creta, "chalk") geologic period.: Smokeybjb, CC BY SA 3.0, via Wikimedia Commons @ https://commons.wikimedia.org/wiki/File:Pakasuchus_dentition.svg

Pakasuchus kapilimai: an extinct, warm-blooded ancestor of saltwater crocodiles (Crocodilus porosus)
life restoration of Pakasuchus kapilimai: Smokeybjb, CC BY SA 3.0, via Wikimedia Commons @ https://commons.wikimedia.org/wiki/File:Pakasuchus.jpg

"Pakasuchus" (1:10)
Uploaded August 8, 2010, by tyrantslayer24 to YouTube ~ URL: https://www.youtube.com/watch?v=DzJ3lxtJKFs

"Pakasuchus teeth" (0:33)
Uploaded August 3, 2010, by edyong209 to YouTube~ URL: https://www.youtube.com/watch?v=u2u0lUfsR8E

Pakasuchus kapilimai's homeland: southeastern Tanzania's Rukwa Rift Basin contains Lake Rukwa, watery homeland of huge populations of modern-day crocodiles.
Lake Rukwa, in Tanzania, as seen from space: NASA World Wind, Public Domain, via Wikimedia Commons @ https://commons.wikimedia.org/wiki/File:Lake_Rukwa.png

 

"Pakasuchus teeth" (0:33)

Uploaded August 3, 2010, by edyong209 to YouTube~ URL: https://www.youtube.com/watch?v=u2u0lUfsR8E

Pakasuchus teeth

Sources Consulted

 

Alonso, Juan Carlos, and Gregory S. Paul. 2015. The Early Cretaceous: Notes, Drawings, and Observations from Prehistory. Lake Forest, CA: Walter Foster Jr., imprint of Quarto Publishing Group USA Inc.

Bonner, John. 4 August 2010. “Tooth to tail oddities in ancient croc.” Nature > News. 

  • Available @ http://www.nature.com/news/2010/100804/full/news.2010.389.html

Kaufman, Gabriel. 2007. Saltwater Crocodile: The World’s Biggest Reptile. New York, NY: Bearport Publishing. 

O’Connor, Patrick M.; Sertich, Joseph J.W.; Stevens, Nancy J.; Roberts, Eric M.; Gottfried, Michael D.; Hieronymus, Tobin L.; Jinnah, Zubair A.; Ridgely, Ryan; Ngasala, Sifa E.; Temba, Jesuit. The evolution of mammal-like crocodyliforms in the Cretaceous Period of Gondwana. Nature 466, 748–751 (5 August 2010). doi:10.1038/nature09061

  • Available @ http://www.nature.com/articles/nature09061.epdf?referrer_access_token=k9ok9Tg2d8jeus4t_S_1l9RgN0jAjWel9jnR3ZoTv0NCPgYDsjdK4EyfgsZwD0mC_ohD9FUgWk2wMj4DhJoBU1Mcdz5QqLzGHRu-sDuxtklc4Kj-xwbg_G_TL75ozqdDz9YYNIXDNWql5-fyDn7XScDaqYXhH8cQff-e-Kn8Xk-CQooAHsnixKZj57la9XWg&tracking_referrer=www.nature.com

Ohio University. 4 August 2010. “Mammal-like crocodile fossil found in East Africa, scientists report.” ScienceDaily > Releases.

  • Available @ http://www.sciencedaily.com/releases/2010/08/100804133352.htm

Strauss, Bob. "10 prehistoric crocodiles everyone should know." About.com > About Education > Dinosaurs > Prehistoric Animals.

  • Available @ http://dinosaurs.about.com/od/otherprehistoriclife/ss/10-Prehistoric-Crocodiles-Everyone-Should-Know.htm#step1

Switek, Brian. 6 August 2010. “Pakasuchus: the croc that ate like a mammal. Smithsonian > Science > Nature.

  • Available @ http://www.smithsonianmag.com/science-nature/pakasuchus-the-croc-that-ate-like-a-mammal-77899607/?no-ist 

Than, Ker. 5 August 2010. “Pictures: Ancient ‘Cat Crocodile’ Discovered.” Illustration Courtesy Zina Deretsky, National Science Foundation. National Geographic > News > Photogalleries.

  • Available @ http://news.nationalgeographic.com/news/2010/08/photogalleries/100804-crocodile-chew-fossil-science-mammal-teeth-pictures/

"These Crocs Are Made for Biting: The Discovery." National Science Foundation > News > Special Reports.

  • Available via NSF @ http://www.nsf.gov/news/special_reports/crocs/discovery.jsp

"These Crocs Are Made for Biting Fact Sheet: Pakasuchus kapilimai." National Science Foundation > News > Special Reports.

  • Available via NSF @ http://www.nsf.gov/news/special_reports/crocs/fact.jsp

"These Crocs Were Made for Chewing?" 2010. National Science Foundation > News. Press Release 10-134 (August 4, 2010).

  • Available via NSF @ http://www.nsf.gov/news/news_summ.jsp?org=NSF&cntn_id=117392

Witton, Mark. 3 August 2010. “Pakasuchus: an ancient crocodile and mammal wannabe.” Discover Magazine > Photos.

  • Available @ http://discovermagazine.com/galleries/zen-photo/p/pakasuchus

Yong, Ed. 4 August 2010. "Pakasuchus -- the crocodile that’s trying to be a mammal." NGM.com > Phenomena, A science salon hosted by National Geographic Magazine.

  • Available @ http://phenomena.nationalgeographic.com/2010/08/04/pakasuchus-the-crocodile-thats-trying-to-be-a-mammal/

Zielinski, Sarah. 6 August 2010. “A cat-like crocodile from the Cretaceous.” Smithsonian > Science > Nature.

  • Available @ http://www.smithsonianmag.com/science-nature/a-cat-like-crocodile-from-the-cretaceous-32854440/

 

Pakasuchus kapilimai's homeland: southeastern Tanzania's Rukwa Rift Basin contains Lake Rukwa, watery homeland of huge populations of modern-day crocodiles.

Lake Rukwa, in Tanzania, as seen from space
Lake Rukwa, in Tanzania, as seen from space — NASA World Wind, Public Domain, via Wikimedia Commons
the end which is also the beginning
the end which is also the beginning — OpenClips, Public Domain, via Pixabay

Ancient Earth Journal: The Early Cretaceous: Notes, Drawings, and Observations from Prehistory by Juan Carlos Alonso and Gregory S. Paul ~ Available via Amazon

Cretaceous geologic period

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Me and my purrfectly purrfect Maine coon kittycat, Augusta "Gusty" Sunshine

DerdriuMarriner, All Rights Reserved

Updated September 11, 2026

Comments

DerdriuMarrinerJan 28, 2025

Lego toys perhaps cooperate more than plush toys with my consideration of country and world gardens and menageries.

Drawing together cat-crocodile allies, ambiances, predators and prey demands something durable like lego components for outside gardens and menageries and something lasting perhaps three, three-plus seasons for indoor gardens and menageries.

Wouldn't indoor and outdoor workings warrant wending around water features?
Potable water woos us all, animal and human and plant sentients, throughout all time from most anciently to most modernly!

DerdriuMarrinerJan 27, 2025

Cat crocodiles develop delightfully into lego and plush toys, don't they?

Such educationally entertaining, entertainingly educational fun to figure cat crocodile allies, habitats, predators and prey!

Such a toy environment might match nicely with a sign "Caution: Time dimension crossing!), beyond which muster lego and plush toys about modern feline sentients and their allies, ambiances, predators (;-{) and prey, mightn't it?

DerdriuMarrinerOct 30, 2024

The computer crashed before I could communicate another component to my comment contained in the box immediately below.

Cat crocodiles with armor-less bodies and armor-full tails intrigue me.
Their tails and their whiskers let feline sentients locate themselves within their environments.

Might cat crocodiles have been as savvy about how they move difficult-ly or easily through their surroundings as the tailed, whiskered feline sentient 99.9-plus and recurring percent correct in where others begin and where they end, where they locate and where they will land?

DerdriuMarrinerOct 4, 2024

The last paragraph to the first subheading, Cat crocodiles exhibit mammal-like mouthparts for piecemeal chews, advises us of "diets of insects, primitive mammals, small lizards."

Sometimes my feline sentients appear to play non-fatal games with insects such as the brown marmorateds.

Might that playfulness perhaps assure them of insect attractiveness as appetizing edibles to alligator ancestors acknowledged as catlike-headed, limbed, lined but not of feline ancestors?

Might feline sentients be mewing, "You're safe with me! You're safe with me!"?

DerdriuMarrinerAug 1, 2015

CruiseReady, Cat crocodiles weren't apex predators so they were part of the everybody-gets-eaten chain in ancient food webs. It's a bit gruesomely intriguing to ponder what their meat tasted like, as possibly more mammalian -- since they were warm-blooded -- than reptilian.

CruiseReadyJul 31, 2015

What amazing creature this must have been... small as the baby gatprs or crocs that we know today, yet with some attributes we don't associate with them at all. Wonder if they were ever prey to their larger cousins?

DerdriuMarrinerJul 14, 2015

happynutritionist, Thank you for liking the images and information on cat crocodiles! Me too, I completely understand the feline comparisons in terms of head, limbs, and lines. But it indeed would be a scary situation for a cat crocodile to expect to be petted, sit on laps, or...swish that armored tail!

DerdriuMarrinerJul 14, 2015

blackspanielgallery, Yes, it's amazing about the archaeological traces that extinction leaves! It must be so rewarding to be one of the artists whose job it is to draw the reconstructions from the available evidence.

happynutritionistJul 9, 2015

I can see why it would be said that this crocodile is catlike, the picture renderings have a posture that looks like a cat, though not one that would be pleasant to pet or have sitting on the lap. Beautifully done page.

blackspanielgalleryJul 7, 2015

It is amazing how much is becoming known of extinct animal.
DerdriuMarriner
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